There is provided a game system and a puzzle game program capable of changing displayed states of puzzle elements depending on a tilt of the puzzle game apparatus, such that a high level of strategy-making is required and drastic game developments and excitement are realized, as well as a storage medium having the puzzle game program stored therein. A tilt detection section is provided in connection with the game apparatus (or a controller therefor) for determining a tilt direction. The puzzle elements displayed on a display screen are moved in a tilt direction of the game apparatus.