An agricultural combine having a supporting structure that is driven by ground engaging wheels at a harvesting speed by a propulsion assembly. The combine is also provided with crop processing assemblies for processing a harvested crop. A feederhouse directs the harvested crop material to the crop processing assemblies. A feederhouse force sensor senses the force exerted by the harvested crop material as it passes through the feederhouse. The feederhouse force signal is directly related to crop throughput. An operator control in the operator's cab of the combine provides a loss rate signal that is also directed to the electronic controller. The electronic controller converts the loss rate signal into a desired feederhouse force signal. The electronic controller regulates the forward speed or harvesting speed of the combine so that the desired feederhouse force signal is equal to the actual feederhouse force signal. The harvesting speed regulation can be modified or overridden by moisture signals from a moisture sensor located in the feederhouse for sensing the moisture content of harvested crop.